生態圈的范圍和生態學本身一樣廣泛。該雜志歡迎來自生態科學各個子學科的投稿，以及與生態學相關的跨學科研究。該雜志的目標是提供一種快速出版的、在線的、開放獲取的歐空局其他期刊的替代品，同時保持歐空局出版物所享有的嚴格的同行評審標準。

The scope of Ecosphere is as broad as the science of ecology itself. The journal welcomes submissions from all sub-disciplines of ecological science, as well as interdisciplinary studies relating to ecology. The journal's goal is to provide a rapid-publication, online-only, open-access alternative to ESA's other journals, while maintaining the rigorous standards of peer review for which ESA publications are renowned.